Faculty Directory

Ashley Dowling

Ashley Dowling

Associate Professor

Diversity, Evolution, Systematics

Director Experiment Station


Phone: 479-575-2451

Fax: (479) 575-2452

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  • B.S., Ecology and Evolutionary Biology, University of Arizona, Tucson, AZ
  • Ph.D., Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 

Current Students

  • Ray Fisher (PhD)
  • Whitney Nelson (PhD)
  • Stephen Robertson(PhD)
  • Alexis Hilario (MS)
  • Colton Zirkle (MS)
  • Phillip Stephenson (MS, coadvised through Biology)

ENTO 3011: Introduction to Insect Identification Lab

ENTO 3013: Introduction to Entomology

Select Publications since 2015 (* indicates graduate student author)

Skvarla, M.J.*, Evans, G.E., Dowling, A.P.G. In press. Clinical report of Otobius megnini (Acari: Parasitiformes: Argasidae) from a domestic cat (Felis silvestris catus) and first record from Northwest Arkansas.

Skvarla, M.J., Fisher, D.M., Schnepp, K.E., Dowling, A.P.G. 2015. Terrestrial arthropods of Steel Creek, Buffalo National River, Arkansas. I. Select beetles (Coleoptera: Buprestidae, Carabidae, Cerambycidae, Curculionoidea excluding Scolytinae). Biodiversity Data Journal 3: e6832. doi:  10.3897/BDJ.3.e6832

Skvarla, M.J.*, Bertone, M.A., Fisher, J.R., Dowling, A.P.G. 2015. New information about the cypress weevil, Eudociminus mannerheimii (Boheman, 1836) (Coleoptera: Curculionidae: Molytinae): redescription, range expansion, new host records, and a report as a possible causative agent of tree mortality. Coleopterists Bulletin 69(4): 751-757

Skvarla, M.J.*, Tripodi, A.*, Szalanski, A., and Dowling, A.P.G. 2015. New records of Orussus minutus Middlekauff, 1983 (Hymenoptera: Orussidae) represent a significant western range expansion. Biodiversity Data Journal 3: e5793. doi: 10.3897/BDJ.3.e5793

Fisher, J.R.*, Fisher, D.M., Nelson, W.A.*, O’Neill, J.C.*, Skvarla, M.J., Radwell, A.J., Ochoa, R., Bauchan, G., and Dowling, A.P.G. 2015. An integrative description of Torrenticola trimaculata sp. nov. (Parasitengona: Torrenticolidae), a three-spotted, riffle-dwelling mite from eastern North America: morphology, phylogenetics, and taxonomic history of the genus. Acarologia. 55(1): 71-116

Dowling, A.P.G. 2015. The Evolution of parasitism and host associations in mites. In: Morand, S., Krasnov, B.R. & Littlewood, D.T.J. (eds). Parasite Diversity and Diversification - Evolutionary Ecology Meets Phylogenetics. Cambridge University Press.          pgs. 265-288.

Fisher, J.R.*, Fisher, D.M., Skvarla, M.J.*, and Dowling, A.P.G. 2014. Pre-ballooning in Ummidia Thorell, 1875 (Araneae: Ctenizidae) from the Interior Highlands, USA: second account from the region and review of mygalomorph ballooning. J. Arachnology. 42(3): 318-321

Skvarla, M.J.*, Larson, J.L., Dowling, A.P.G. 2014. Pitfalls and preservatives: A review. J. Entomol. Soc. Ontario. 145: 15-43.

Skvarla, M.J.* and Dowling, A.P.G. 2014. First report of gynandromorphism in Temnothorax curvispinosus (Mayr, 1886) (Hymenoptera: Formicidae).  Proc. Entomol. Soc. Washington. 116(3): 349-353.

Skvarla, M.J.*, Hartshorn, J.A.*, Dowling, A.P.G. 2014. Report on a large collection of Merope tuber Newman, 1838 (Mecoptera: Meropeidae) from Arkansas with notes on collection technique, sex ratio, and male clasper size. Psyche 2014, Article ID 530757, 6 pages.

Wiedenmann, R.N., Dowling, A.P.G., and Barnes, J.K. 2014. What is the value of your insect collection? Amer. Entomologist 60(2): 101-104

Skvarla, M.J.*, Fisher, J.R.*, and Dowling, A.P.G. 2014. A review of Cunaxidae (Acariformes, Trombidiformes): Histories and diagnoses of subfamilies and genera, keys to world species, and some new locality records. Zookeys 418:1-103

Skvarla, M.J.*, Fisher, J.R.*, and Dowling, A.P.G. 2013. On some mites (Acari: Prostigmata) from the Interior Highlands: descriptions of the male, immature stages, and female reproductive systems of Pseudocheylus americanus (Ewing, 1909) and some new state records for Arkansas.  Zootaxa, 3641(4): 401–419.

Paré, J.A. and Dowling, A.P.G. 2012. An overview of acariasis in captive invertebrates. J. Zoo Wildlife Med. 43: 703-714

Beard, J.J., Ochoa, R., Bauchan, G.R., Welbourn, W.C., Pooley, C. and Dowling, A.P.G. 2012. External mouthpart morphology in the Tenuipalpidae (Tetranychoidea): Raoiella a case study. Exp. Appl. Acarol. 57: 227-255

Dowling, A.P.G., Ochoa, R., Beard, J.J., Welbourn, W.C., and Ueckermann, E.A. 2012.  Phylogenetic investigation of the genus Raoiella (Prostigmata: Tenuipalpidae): Diversity, distribution, and world invasions. Exp. Appl. Acarol.57: 257-269

Skvarla, M.J.* and Dowling, A.P.G. 2012. Some new armascirine cunaxids (Acari: Prostigmata: Cunaxidae) from the Eastern United States. Zootaxa 3194: 1-34

Radwell, A.J., Dowling, A.P.G., Smith, I.M., and Kaliki¥, V. 2011. Kongsbergia robisoni, n. sp. (Acari: Hydrachnidiae: Aturidae) from the interior highlands of North America based on morphology and molecular genetic analysis. Int. J. Acarol. 37: 194-205

Skvarla, M.J.*, Fisher, J.R.*, and Dowling, A.P.G. 2011. A new species of Neoscirula (Acari: Prostigmata: Cunaxidae) from the Ozark Highlands. Acarologia 51(3): 283-293

Beaulieu, F., Dowling, A.P.G., Klompen, H., De Moraes, G.J., and Walter, D.E. 2011. Superorder Parasitiformes Reuter, 1909. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 123-128.

Fisher, J.R.*, Skvarla, M.J.*, Bauchan, G.R., Ochoa, R., and Dowling, A.P.G. 2011. Trachymolgus purpureus sp. nov., an armored snout mite (Acari: Bdellidae) from the Ozark Highlands: morphology, development, and key to Trachymolgus Berlese. ZooKeys 125: 1-34.

Dowling, A.P.G., Bauchan, G.R., Beard, J.J., and Ochoa, R. 2010. Scanning electron microscopy vouchers and genomic data from an individual specimen: maximizing the utility of delicate and rare specimens. Acarologia 50: 479-485.

Fisher, J.R.* and Dowling, A.P.G. 2010. Modern methods and technology for doing classical taxonomy. Acarologia 50: 395-409.

Dowling, A.P.G. and OConnor, B.M. 2010. Phylogenetic relationships within the suborder Dermanyssina (Acari: Parasitiformes) and a test of dermanyssoid monophyly. Int. J. Acarol. 36: 299-312.

Roy, L.*, Dowling, A.P.G., Chauve, C.M. and Buronfosse, T. 2010. Diversity of phylogenetic information according to the locus and the taxonomic level: an example from a parasitic mesostigmatid mite genus. Int. J. Mol. Sci. 11: 1704-1734.

Dowling, A.P.G. and OConnor, B.M. 2010. Phylogeny of Dermanyssoidea (Acari: Parasitiformes) suggests multiple origins of parasitism. Acarologia 50: 113-129.

Associate Professor, 2013-present, Department of Entomology, University of Arkansas

Assistant Professor, 2008-2013, Department of Entomology, University of Arkansas

Postdoctoral Scholar, 2005-2007, Department of Entomology, University of Kentucky

Instructor Ohio State Summer Acarology Program, 2010, 2011, 2012, 2014, 2016

Gamma Sigma Delta Teaching Award of Excellence, 2014

University of Arkansas Outstanding Mentor Award, 2014