EDUCATION:

• B. A., Entomology, San José State University, San José, CA, 1967
• Ph.D., Entomology, University of California, Berkeley, Berkeley, CA, 1974

GENERAL AREAS OF INTEREST:

Forest Entomology, Insect Ecology, Biological Control

SPECIFIC AREAS OF RESEARCH:

• Red Oak Borer: Population dynamics, distribution and geospatial analysis.
• Southern Pine Beetle: Biological control through enhancement of parasitoid effectiveness.
• Southern Pine Beetle: Population and infestation growth modeling.
• Gypsy Moth: Effects of Bt eradication sprays on nontarget canopy arthropods and their bird predators.

SELECTED PUBLICATIONS:

• Aquino, L.D., Tullis, J.A., and Stephen, F.M.. 2008.Modeling red oak borer, Enaphalodes rufulus(Haldeman), damage using in situ and ancillary landscape data. Forest Ecology and Management 255 (2008) 931–939.
• Haavik, L.J., Stephen, F.M., Fierke, M.K., Salisbury, V.B., Leavitt, S.W., and Billings, S.A.. 2008.Dendrochronological parameters of northern red oak (Quercus rubra L.(Fagaceae)) infested with red oak borer (Enaphalodes rufulus (Haldeman)(Coleoptera: Cerambycidae)). Forest Ecology and Management 255 (2008)1501–1509.
• Fierke, M.K., and Stephen, F.M. 2007. Red Oak Borer (Coleoptera: Cerambycidae) Flight Trapping In The Ozark National Forest, Arkansas. Florida Entomologist 90(3):488-494.
• Fierke, M.K., Kelley, M.B., Stephen, F.M. 2007. Site and stand variables influencing red oak borer, Enaphalodes rufulus (Coleoptera: Cerambycidae), population densities andtree mortality. Forest Ecology and Management 247 (2007) 227–236.
• Crook, D.J., M.K. Fierke, A. Mauromoustakos, D.L. Kinney, F.M. Stephen. 2007. Optimization of sampling methods for within-tree populations of red oak borer, Enaphalodes rufulus (Haldeman) (Coleoptera: Cerambycidae). Environ. Entomol. 36:589-594.
• Fierke, M.K., Kinney, D.L., Salisbury, V.B., Crook, D.J., and Stephen, F.M. 2005. A rapid estimation procedure for within-tree populations of red oak borer (Coleoptera: Cerambycidae). Forest Ecology and Management 215 (2005) 163–168.
• M. K. Fierke; D. L. Kinney; V. B. Salisbury; D. J. Crook; F. M. Stephen. (2005) Development and Comparison of Intensive and Extensive Sampling Methods and Preliminary Within-Tree Population Estimates of Red Oak Borer (Coleoptera: Cerambycidae) in the Ozark Mountains of Arkansas. Environmental Entomology. 34(1):184 - 192.
• Stephen, F. M., V. B. Salisbury and F. L. Oliveria. 2003. Red oak borer, Enaphalodes rufulus (Coleoptera: Cerambycidae), in the Ozark Mountains of Arkansa, USA: an unexpected and remarkable forest disturbance. Integrated Pest Management Reviews.6:247-252.
• Stephen, F. M., V. B. Salisbury, W. F. Limp & D. J.Crook. (2002) Red oak borer, Enaphalodes rufulus (Coleoptera: Cerambycidae), a Remarkable Epidemic of Oak Mortality In The Interior Highland Forests. AFRC-CES sponsored symposium in Little Rock, AR May 23, 2002.
• Dodds, K. J., C. Graber and F. M. Stephen. 2001. Facultative intraguild predation by larval Cerambycidae (Coleoptera), Monochamus carolinensis, on bark beetle larvae (Coleoptera: Scolytidae). Environ. Entomol. 30(1): 17-22.
• Stephen, F. M. and J. C. Gregoire. 2001. Introduction and establishment of exotic bark beetles In. Risks of Exotic Forest Pests and their Impact on Trade. April 19-29, 2001 http://www.apsnet.org/meetings/ExoticPests/top.htm . Amer Phytopath. Assn., USDA Forest Service, et. al.
• Stephen, F. M. and L. E. Browne. 2000. Application of Eliminade TM parasitoid food to boles and crowns of pines ( Pinaceae ) infested with Dendroctonus frontalis (Coleoptera: Scolytidae). Can. Entomol. 132:983-985.
• Dodds, K. J. and F. M. Stephen. 2000. Partial age-specific life tables for Monochamus titillator in Dendroctonus frontalis-infested loblolly pines. Entomologia Experimentalis et Applicata 97:331-338.
• Mathews, P. L. and F. M. Stephen. 1999. Impacts of an artificial diet and varied environmental conditions on longevity of Coeloides pissodis (Hymenoptera: Braconidae), a parasitoid of Dendroctonus frontalis (Coleoptera: Scolytidae). Environ. Entomol. 28: 729-734.
• Stephen, F. M., M. P. Lih and L. E. Browne. 1997. Augmentation of Dendroctonus frontalis parasitoid effectiveness by artificial diet. In J. C. Gregoire, A. M. Liebhold, F. M. Stephen, K. R. Day and S. M. Salom, editors. Proceedings: Integrating cultural tactics into the management of bark beetle and reforestation pests. Gen. Tech. Rep. NE- . Radnor, PA. USDA Forest Service. NE For. Exp. Sta. 
• Lih, M. P. and F. M. Stephen. 1995. Relationship of host tree diameter to within-tree southern pine beetle (Coleoptera: Scolytidae) population dynamics. Environ. Entomol. 25:736-742. 
• Lih, M. P., F. M. Stephen, K. G. Smith, L. R. Nagy, G. W. Wallis, and L. C. Thompson. 1995. Effects of a gypsy moth eradication program on nontarget forest canopy insects and their bird predators. Proc. Annual Gypsy Moth Review, Portland, OR, Oct. 30-Nov. 2, 1994. 
• Stephen, F. M. 1995. Potential for suppressing southern pine beetle populations by enhancing effectiveness of their hymenopteran parasitoids. pp. 226-240. In Behavior, population dynamics and control of forest insects. In F. P. Hain, S. M. Salom, W. F. Ravlin, T. L. Payne and K. F. Raffa, editors. Proceedings: Behavior, population dynamics and control of forest insects. Ohio State University, Ohio Agricultural Research and Development Center: Wooster, Ohio. 
• Lorio, P. L. Jr., Stephen, F. M. and T. D. Paine. 1995. Environment and ontogeny modify loblolly pine response to induced acute water deficits and bark beetle attack. For. Ecology and Manag. 73:97-110.
• Jones, J. M. and F. M. Stephen. 1994 Effect of temperature on development of hymenopterous parasitoids of Dendroctonus frontalis (Coleoptera: Scolytidae). Environ. Entomol. 23:457-463. 
• Williams, K. S., K. G. Smith and F. M. Stephen. 1993. Emergence of 13-year periodical cicadas (Cicadidae: Magicicada): phenology, mortality, and predator satiation. Ecology 74:1143-1152.  

COURSES TAUGHT:

• ENTO 4053/4050L : Insect Ecology / Insect Ecology Laboratory (Website )

GRADUATE STUDENTS AND STAFF:

• John Riggins - Graduate Student (Ph.D.)
• Larry Galligan - Research Specialist
• Robin Verble - Graduate Student (M.S.)
• Laurel Haavik - Graduate Student (Ph.D.)

F. M. Stephen Department of Entomology - 319 Agriculture Building University of Arkansas, Fayetteville Fayetteville, Arkansas 72701 Dr. Stephen: (479) 575-3404 Facsimile: (479) 575-2452

E-Mail: Dr. Stephen Forest Entomology Site Short CV